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" ... Thus, this peculiar bird is the product of male and female fraternal twin embryos, resulting from two different ova fertilized by two different sperms. Somewhere between the 2-cell and the 64-cell stage of development, these male and female embryos that had been developing alongside each other inside the same eggshell ceased to develop separately and fused into just one embryo. This bizarre bird is that embryo -- all grown up. It exemplifies an unusual phenomenon, a fascinating developmental mistake, known in scientific circles as a bilateral gynandromorph, and amongst veterinarians and pet bird breeders -- and even by some bird watchers -- as a half-sider. Because northern cardinals are a sexually dimorphic species, where males are scarlet and females are tan, it was easy to see that this peculiar bird is both male and female. (Recognizing a bilateral gynandromorph is almost impossible when looking at species where males and females appear identical.) ... "
" ... Whilst this bottleneck on average reduces the mutation load of the offspring below that of the mother, some proportion of mutant mitochondria can still survive. And if the specific cell lineage that gave rise to the mother’s ova happened to have a high mutant load, then her offspring may still have a higher load than she does. ... "